Laughter is a pleasant physical reaction and emotion consisting usually of rhythmical, often audible contractions of the diaphragm and other parts of the respiratory system. It is a response to certain external or internal stimuli. Laughter can rise from such activities as being tickled, or from humorous stories or thoughts. Most commonly, it is considered an auditory expression of a number of positive emotional states, such as joy, mirth, happiness, or relief. On some occasions, however, it may be caused by contrary emotional states such as embarrassment, surprise, or confusion such as nervous laughter or courtesy laugh. Age, gender, education, language, and culture are all indicators as to whether a person will experience laughter in a given situation. Other than humans, some other species of primate show laughter-like vocalizations in response to physical contact such as wrestling, play chasing or tickling.